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In both snapdragon and Arabidopisis, the floral B-function, which specifies petal and stamen development, is embedded in a heterodimer consisting of one DEFICIENS/APETALA3-like and one GLOBOSA/PISTILLATA-like MADS box protein. Vandenbussche et al. (pages 741-754) demonstrate that gene duplications in both the DEF/AP3 and GLO/PI lineages in Petunia have led to a functional diversification of their respective members, which is reflected by partner specificity and whorl-specific functions among these proteins. The image shows a Petunia phglo1 phglo2 double mutant displaying a full homeotic conversion of petals to sepals and of stamens to carpels in the flower, a phenotype very similar to single B-function mutants in Arabidopsis and snapdragon.